PROSITE documentation PDOC00803

ATP-dependent serine proteases, lon family, serine active site

Description:

The following proteins belongs to a family of proteases which are dependent on the hydrolysis of ATP for their activity and which have a serine in their active site:

  • Bacterial ATP-dependent proteases [1,2]. The prototype of those bacterial enzymes is the Escherichia coli La protease (EC 3.4.21.53) (gene lon). La is capable of hydrolyzing large proteins; it degrades short-lived regulatory (such as rcsA and sulA) and abnormal proteins. It is a cytoplasmic protein of 87 Kd that associates as an homotetramer. Its proteolytic activity is stimulated by single-stranded DNA.
  • Eukaryotic mitochondrial matrix proteases [3,4]. The prototype of these enzymes is the yeast PIM1 protease. It is a mitochondrial matrix protein of 120 Kd that associated as an homohexamer. It catalyzes the initial step of mitochondrial protein degradation.
  • Haemophilus influenzae lon-B (HI1324), a protein which does not contain the ATP-binding domain, but possess a slightly divergent form of the catalytic domain.

The region around the serine residue involved in the catalytic mechanism [5] is perfectly conserved and can be used as a signature pattern.

Note:

Proteins belonging to this family also contain a copy of the ATP/GTP- binding motif 'A' (P-loop) (see <PDOC00017>).

Note:

These proteins belong to family S16 in the classification of peptidases [6,E1].

Last update:

November 1995 / First entry.

Technical section:

PROSITE method (with tools and information) covered by this documentation:

LON_SER, PS01046ATP-dependent serine proteases, lon family, serine active site  (PATTERN)
Consensus pattern: D-G-[PD]-S-A-[GS]-[LIVMCA]-[TA]-[LIVM]
S is the active site residue
Sequences known to belong to this class detected by the pattern: ALL
Other sequence(s) detected in Swiss-Prot: NONE.
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Matching PDB structures: 2X36 [ALL]

References:

1 AuthorsThomas C.D., Modha J., Razzaq T.M., Cullis P.M., Rivett A.J.
TitleControlled high-level expression of the lon gene of Escherichia coli allows overproduction of Lon protease.
SourceGene 136:237-242(1993).
PubMed ID8294008
2 AuthorsTojo N., Inouye S., Komano T.
TitleThe lonD gene is homologous to the lon gene encoding an ATP-dependent protease and is essential for the development of Myxococcus xanthus.
SourceJ. Bacteriol. 175:4545-4549(1993).
PubMed ID8331083
3 Authorsvan Dyck L., Pearce D.A., Sherman F.
SourceJ. Biol. Chem. 269:238-242(1994).
4 AuthorsWang N., Gottesman S., Willingham M.C., Gottesman M.M., Maurizi M.R.
TitleA human mitochondrial ATP-dependent protease that is highly homologous to bacterial Lon protease.
SourceProc. Natl. Acad. Sci. U.S.A. 90:11247-11251(1993).
PubMed ID8248235
5 AuthorsFischer H., Glockshuber R.
TitleATP hydrolysis is not stoichiometrically linked with proteolysis in the ATP-dependent protease La from Escherichia coli.
SourceJ. Biol. Chem. 268:22502-22507(1993).
PubMed ID8226758
6 AuthorsRawlings N.D., Barrett A.J.
TitleFamilies of serine peptidases.
SourceMethods Enzymol. 244:19-61(1994).
PubMed ID7845208
E1
Sourcehttp://www.expasy.org/cgi-bin/lists?peptidas.txt

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