PROSITE documentation PDOC00856

RNA polymerases N / 8 Kd subunits signature

Description:

In eukaryotes, there are three different forms of DNA-dependent RNA polymerases (EC 2.7.7.6) transcribing different sets of genes. Each class of RNA polymerase is an assemblage of ten to twelve different polypeptides. In archaebacteria, there is generally a single form of RNA polymerase which also consist of an oligomeric assemblage of 10 to 13 polypeptides.

Archaebacterial subunit N (gene rpoN) [1] is a small protein of about 8 Kd, it is evolutionary related [2] to a 8.3 Kd component shared by all three forms of eukaryotic RNA polymerases (gene RPB10 in yeast and POLR2J in mammals) as well as to African swine fever virus protein CP80R [3].

As a signature pattern we selected a conserved region which is located at the N-terminal extremity of these polymerase subunits; this region contains two cysteines that binds a zinc ion [4].

Last update:

May 2004 / Text revised.

Technical section:

PROSITE method (with tools and information) covered by this documentation:

RNA_POL_N_8KD, PS01112RNA polymerases N / 8 Kd subunits signature  (PATTERN)
Consensus pattern: [LIVMFD]-[LIVMFP]-P-[LIVM]-x-C-[FL]-[ST]-C-G
The 2 C's are zinc ligands
Sequences known to belong to this class detected by the pattern: ALL
Other sequence(s) detected in Swiss-Prot: NONE
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Matching PDB structures: 1EF4 1I3Q 1I50 1I6H ... [ALL]

References:

1 AuthorsLanger D., Hain J., Thuriaux P., Zillig W.
TitleTranscription in archaea: similarity to that in eucarya.
SourceProc. Natl. Acad. Sci. U.S.A. 92:5768-5772(1995).
PubMed ID7597027
2 AuthorsMcKune K., Woychik N.A.
TitleHalobacterial S9 operon contains two genes encoding proteins homologous to subunits shared by eukaryotic RNA polymerases I, II, and III.
SourceJ. Bacteriol. 176:4754-4756(1994).
PubMed ID8045907
3 AuthorsYanez R.J., Rodriguez J.M., Nogal M.L., Yuste L., Enriquez C., Rodriguez J.F., Vinuela E.
TitleAnalysis of the complete nucleotide sequence of African swine fever virus.
SourceVirology 208:249-278(1995).
PubMed ID11831707
4 AuthorsMacKereth C.D., Arrowsmith C.H., Edwards A.M., McIntosh L.P.
TitleZinc-bundle structure of the essential RNA polymerase subunit RPB10 from Methanobacterium thermoautotrophicum.
SourceProc. Natl. Acad. Sci. U.S.A. 97:6316-6321(2000).
PubMed ID10841539
DOI97/12/6316

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