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We are deeply saddened by the passing of Amos Bairoch (1957–2025), the creator of PROSITE. We wish to dedicate our latest paper, published shortly before his death, to him. He will always be a source of inspiration to us.
Our deepest condolences go out to his family and friends, and to all those who had the privilege of working with him. Rest in peace, Amos. Your work will live on long after you are gone.
Amos Bairoch

PROSITE documentation PDOC50890
PUA domain profile


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PURL: https://purl.expasy.org/prosite/documentation/PDOC50890

Description

The PUA domain, which has been named after PseudoUridine synthase and Archaeosine-specific transglycosylase, is a small, compact domain that consists of 78-83 amino acid residues. The PUA domain is widely conserved in eukaryotic and archaeal RNA modification enzymes and is found in:

  • Archaeal and eukaryotic pseudouridine synthases.
  • Archaeosine tRNA-guanine transglycosylases.
  • A family of predicted ATPases that may be involved in RNA modification.
  • A family of predicted archaeal and bacterial rRNA methylases.
  • A conserved family of putative novel eukaryotic translation factors.
  • Glutamate 5-kinases (EC 2.7.2.11).

The PUA domain can be found alone or in association with other domains, such as the glutamate kinase domain (see <PDOC00701>), the pseudouridine synthase catalytic domain, the phosphoadenosine phosphosulfate (PAPS) reductase domain, the methylase domain, the TGT domain, or the SUI1 domain (see <PDOC00862 >). The PUA domain is supposed to bind to RNA molecules with complex folded structures [1].

The PUA domain contains highly conserved motifs that center around stretches of hydrophobic residues. The domain also contains two highly conserved positions occupied by small amino acids (primarily glycines) [1]. The resolution of the crystal structure of archaeosine tRNA-guanine transglycosylase from Pyrococcus horikoshii has shown that the PUA domain consists of two α-helices and six β-strands, and folds into a β-sandwich structure similar to the oligonucleotide-binding (OB)-fold (see <PDB:1IQ8>) [2].

The profile we developed covers the entire PUA domain.

Last update:

November 2002 / First entry.

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Technical section

PROSITE method (with tools and information) covered by this documentation:

PUA, PS50890; PUA domain profile  (MATRIX)


References

1AuthorsAravind L. Koonin E.V.
TitleNovel predicted RNA-binding domains associated with the translation machinery.
SourceJ. Mol. Evol. 48:291-302(1999).
PubMed ID10093218

2AuthorsIshitani R. Nureki O. Fukai S. Kijimoto T. Nameki N. Watanabe M. Kondo H. Sekine M. Okada N. Nishimura S. Yokoyama S.
TitleCrystal structure of archaeosine tRNA-guanine transglycosylase.
SourceJ. Mol. Biol. 318:665-677(2002).
PubMed ID12054814
DOI10.1016/S0022-2836(02)00090-6



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