PROSITE logo
Black ribbon
We are deeply saddened by the passing of Amos Bairoch (1957–2025), the creator of PROSITE. We wish to dedicate our latest paper, published shortly before his death, to him. He will always be a source of inspiration to us.
Our deepest condolences go out to his family and friends, and to all those who had the privilege of working with him. Rest in peace, Amos. Your work will live on long after you are gone.
Amos Bairoch

PROSITE documentation PDOC51262
COS domain profile


View entry in original PROSITE document format
View entry in raw text format (no links)
PURL: https://purl.expasy.org/prosite/documentation/PDOC51262

Description

N-terminal RING finger (see <PDOC00449>)/B-box (see <PDOC50119>)/coiled coil (RBCC) or tripartite motif (TRIM) proteins, which are found in metazoan, are involved in a vast array of intracellular functions. They appear to function as part of large protein complexes and possess ubiquitin-protein isopeptide ligase activity. The following RBCC proteins contain an ~60-residue COS (C-terminal subgroup one signature) domain, which is also found in a distantly related non-RBCC microtubule-binding protein, GLFND:

  • Vertebrate MID1 and MID2, which associate with microtubules through homo- and heterodimerization.
  • Animal TRIM9, which plays a regulatory role in synaptic vesicle exocytosis.
  • Mammalian TRIM nine-like (TNL).
  • Mammalian TRIM36, which could play a regulatory role in exocytosis of the sperm vesicle.
  • Mammalian tripartite, fibronectin type III and C-terminal B30.2/SPRY (TRIFIC).
  • Mammalian muscle-specific RING finger (MURF) family. MURF proteins have an ability to form both homo- and heterodimers with each other and to associate with the microtubule cytoskeleton.

In addition to RBCC, the COS domain is also found in association with B30.2/SPRY (see <PDOC50188>) or fibronectin type-III (FN3) (see <PDOC50853>) domains.

The COS domain is predicted to consist of two α-helical coils [1].

The profile we developed covers the entire COS domain.

Last update:

November 2006 / First entry.

-------------------------------------------------------------------------------


Technical section

PROSITE method (with tools and information) covered by this documentation:

COS, PS51262; COS domain profile  (MATRIX)


Reference

1AuthorsShort K.M. Cox T.C.
TitleSubclassification of the RBCC/TRIM superfamily reveals a novel motif necessary for microtubule binding.
SourceJ. Biol. Chem. 281:8970-8980(2006).
PubMed ID16434393
DOI10.1074/jbc.M512755200



PROSITE is copyrighted by the SIB Swiss Institute of Bioinformatics and distributed under the Creative Commons Attribution-NonCommercial-NoDerivatives (CC BY-NC-ND 4.0) License, see prosite_license.html.