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We are deeply saddened by the passing of Amos Bairoch (1957–2025), the creator of PROSITE. We wish to dedicate our latest paper, published shortly before his death, to him. He will always be a source of inspiration to us.
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Amos Bairoch

PROSITE documentation PDOC51862
BetaSPN-type cysteine-stabilized alpha/beta (CS-alpha/beta) domain profile


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PURL: https://purl.expasy.org/prosite/documentation/PDOC51862

Description

Scorpion venoms are complex mixture of peptides with a variety of pharmacological functions. These toxin peptides include ion channel modulators, antibacterial peptides, and protease inhibitors. Scorpion toxins targeting various ion channels share a cysteine-stabilized α/β (CS-α/β) structure, which can be divided into two types: short-chain toxins of 23-42 amino acid residues with 3 or 4 disulfide bridges, which are commonly potassium channel blockers, and long-chain toxins of 53-78 amino acids, which are mostly modulators of sodium channels.

In addition to the classic short-chain toxins such as α-KTxs, which are specific for potassium channels, long-chain toxins with unique structure and function, which were named β-KTxs and scorpine-like peptides, were identified from the scorpion families Buthidae, Scorpionidae and Caraboctonidae. Peptides of the βSPN (β-KTxs and scorpines) family, which are 59-75 amino acid residues in length, display various antimicrobial, cytolytic, and potassium channel-blocking activities. βSPN family full-length mature peptides contain two structural domains that confer them bi-functionality: the amphipathic α-helical N-terminal domain has cytolytic or antimicrobial activity, while the three-disulfid bridged C-terminal domain with the consensus CS-α/β motif has K(+) channel blocking activity. Sequence analysis revealed that the βSPN family can be divided into three distinct groups: (1) β-KTx-like peptides from buthids; (2) Scorpine-like peptides from Scorpionidae and Caraboctonidae species, including scorpine, Opiscorpines 1-4, HgeScplp1, HgeScplp2 and Heteroscorpine 1; (3) heterogeneous peptides similar to BmTXKβ of buthids and iurids [1,2,3,4,5].

The βSPN-type CS-α/β domain is more closely related to invertebrate defensins, antimicrobial peptides involved in the innate immune response of several invertebrate groups (see <PDOC00356>), than to the classical scorpion toxins. Its structure shows an α helix along with one β sheet stabilized by three disulfide bridges, folding into a CS-α/β motif (see <PDB:5IPO>). The connectivity between the ordinal numbered cysteines is C1-C4, C2-C5, and C3-C6 [5].

The profile we developed covers the entire βSPN-type CS-α/β domain.

Last update:

April 2018 / First entry.

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Technical section

PROSITE method (with tools and information) covered by this documentation:

BSPN_CSAB, PS51862; BetaSPN-type cysteine-stabilized alpha/beta (CS-alpha/beta) domain profile  (MATRIX)


References

1AuthorsDiego-Garcia E. Schwartz E.F. D'Suze G. Gonzalez S.A.R. Batista C.V.F. Garcia B.I. de la Vega R.C. Possani L.D.
TitleWide phylogenetic distribution of Scorpine and long-chain beta-KTx-like peptides in scorpion venoms: identification of 'orphan' components.
SourcePeptides 28:31-37(2007).
PubMed ID17141373
DOI10.1016/j.peptides.2006.06.012

2AuthorsFeng J. Yu C. Wang M. Li Z. Wu Y. Cao Z. Li W. He X. Han S.
TitleExpression and characterization of a novel scorpine-like peptide Ev37, from the scorpion Euscorpiops validus.
SourceProtein Expr. Purif. 88:127-133(2013).
PubMed ID23262394
DOI10.1016/j.pep.2012.12.004

3AuthorsLuna-Martinez K. Jimenez-Vargas J.M. Possani L.D.
TitleScorpine-Like Peptides. Single Cell Biology 5(2016).
DOI10.4172/2168-9431.1000138

4AuthorsZhu S. Gao B. Aumelas A. del Carmen Rodriguez M. Lanz-Mendoza H. Peigneur S. Diego-Garcia E. Martin-Eauclaire M.-F. Tytgat J. Possani L.D.
TitleMeuTXKbeta1, a scorpion venom-derived two-domain potassium channel toxin-like peptide with cytolytic activity.
SourceBiochim. Biophys. Acta 1804:872-883(2010).
PubMed ID20045493
DOI10.1016/j.bbapap.2009.12.017

5AuthorsFlores-Solis D. Toledano Y. Rodriguez-Lima O. Cano-Sanchez P. Ramirez-Cordero B.E. Landa A. Rodriguez de la Vega R.C. Del Rio-Portilla F.
TitleSolution structure and antiparasitic activity of scorpine-like peptides from Hoffmannihadrurus gertschi.
SourceFEBS Lett. 590:2286-2296(2016).
PubMed ID27314815
DOI10.1002/1873-3468.12255



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