PROSITE logo
Black ribbon
We are deeply saddened by the passing of Amos Bairoch (1957–2025), the creator of PROSITE. We wish to dedicate our latest paper, published shortly before his death, to him. He will always be a source of inspiration to us.
Our deepest condolences go out to his family and friends, and to all those who had the privilege of working with him. Rest in peace, Amos. Your work will live on long after you are gone.
Amos Bairoch

PROSITE documentation PDOC51912
DMAP1-binding domain profile


View entry in original PROSITE document format
View entry in raw text format (no links)
PURL: https://purl.expasy.org/prosite/documentation/PDOC51912

Description

The DNA methyltransferase 1 (DNMT1) is a multifunctional protein that is a critical DNA methyltransferase important for chromatin structure and gene silencing as well as methyltransferase-independent transcriptional repression. DNMT1 is composed of multiple functional domains. The N-terminal regulatory region contains conserved domains, including the DNA methyltransferase-associated protein 1 (DMAP1)-binding domain, the PCNA (proliferating cell nuclear antigen) binding domain (PBD) and the replication foci targeting sequence (RFTS) which control the subnuclear localization of DNMT1. In contrast, the C-terminal portion contains a catalytic domain (see <PDOC51555>). The DMAP1-binding domain is a ~120-amino acid protein-protein interaction module that binds notably DMAP1, a transcriptional co-repressor [1,2,3].

In addition to DNMT1, a DMAP1-binding domain is found in the following proteins:

  • Animal disco-interacting protein 2 (DIP-2), that maintains morphology of mature neurons. DIP-2 consists of a DMAP1-binding domain and two adenylate- forming domains (AFDs) [4,5,6].
  • Animal N-acetylglucosamine-1-phosphotransferase subunits α (GNPTA) and γ (GNPTG), members of a complex that catalyzes the initial step in the formation of the mannose 6-phopsphate targeting signal on newly synthesized lysosomal acid hydrolases. The DMAP1-binding domain mediates the selective binding GlcNAc-1-phosphotranferase to acid hydrolases [7,8].

The DMAP1-binding domain is predicted to adopt a long helix-turn-helix structure that is rich in leucine residues [1].

The profile we developed covers the entire DMAP1-binding domain.

Last update:

January 2020 / First entry.

-------------------------------------------------------------------------------


Technical section

PROSITE method (with tools and information) covered by this documentation:

DMAP1_BIND, PS51912; DMAP1-binding domain profile  (MATRIX)


References

1AuthorsYoder J.A. Yen R.-W. Vertino P.M. Bestor T.H. Baylin S.B.
TitleNew 5' regions of the murine and human genes for DNA (cytosine-5)-methyltransferase.
SourceJ. Biol. Chem. 271:31092-31097(1996).
PubMed ID8940105
DOI10.1074/jbc.271.49.31092

2AuthorsRountree M.R. Bachman K.E. Baylin S.B.
TitleDNMT1 binds HDAC2 and a new co-repressor, DMAP1, to form a complex at replication foci.
SourceNat. Genet. 25:269-277(2000).
PubMed ID10888872
DOI10.1038/77023

3AuthorsMisaki T. Yamaguchi L. Sun J. Orii M. Nishiyama A. Nakanishi M.
TitleThe replication foci targeting sequence (RFTS) of DNMT1 functions as a potent histone H3 binding domain regulated by autoinhibition.
SourceBiochem. Biophys. Res. Commun. 470:741-747(2016).
PubMed ID26774338
DOI10.1016/j.bbrc.2016.01.029

4AuthorsWinnepenninckx B. Debacker K. Ramsay J. Smeets D. Smits A. FitzPatrick D.R. Kooy R.F.
TitleCGG-repeat expansion in the DIP2B gene is associated with the fragile site FRA12A on chromosome 12q13.1.
SourceAm. J. Hum. Genet. 80:221-231(2007).
PubMed ID17236128
DOI10.1086/510800

5AuthorsNitta Y. Yamazaki D. Sugie A. Hiroi M. Tabata T.
TitleDISCO Interacting Protein 2 regulates axonal bifurcation and guidance of Drosophila mushroom body neurons.
SourceDev. Biol. 421:233-244(2017).
PubMed ID27908785
DOI10.1016/j.ydbio.2016.11.015

6AuthorsNoblett N. Wu Z. Ding Z.H. Park S. Roenspies T. Flibotte S. Chisholm A.D. Jin Y. Colavita A.
TitleDIP-2 suppresses ectopic neurite sprouting and axonal regeneration in mature neurons.
SourceJ. Cell. Biol. 218:125-133(2019).
PubMed ID30396999
DOI10.1083/jcb.201804207

7AuthorsQian Y. Flanagan-Steet H. van Meel E. Steet R. Kornfeld S.A.
TitleThe DMAP interaction domain of UDP-GlcNAc:lysosomal enzyme N-acetylglucosamine-1-phosphotransferase is a substrate recognition module.
SourceProc. Natl. Acad. Sci. U. S. A. 110:10246-10251(2013).
PubMed ID23733939
DOI10.1073/pnas.1308453110

8AuthorsQian Y. van Meel E. Flanagan-Steet H. Yox A. Steet R. Kornfeld S.
TitleAnalysis of mucolipidosis II/III GNPTAB missense mutations identifies domains of UDP-GlcNAc:lysosomal enzyme GlcNAc-1-phosphotransferase involved in catalytic function and lysosomal enzyme recognition.
SourceJ. Biol. Chem. 290:3045-3056(2015).
PubMed ID25505245
DOI10.1074/jbc.M114.612507



PROSITE is copyrighted by the SIB Swiss Institute of Bioinformatics and distributed under the Creative Commons Attribution-NonCommercial-NoDerivatives (CC BY-NC-ND 4.0) License, see prosite_license.html.