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We are deeply saddened by the passing of Amos Bairoch (1957–2025), the creator of PROSITE. We wish to dedicate our latest paper, published shortly before his death, to him. He will always be a source of inspiration to us.
Our deepest condolences go out to his family and friends, and to all those who had the privilege of working with him. Rest in peace, Amos. Your work will live on long after you are gone.
Amos Bairoch

PROSITE documentation PDOC51913
HARE-type HTH domain profile


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PURL: https://purl.expasy.org/prosite/documentation/PDOC51913

Description

Human ASXL proteins, orthologs of Drosophila Additional Sex combs, have been implicated in conjunction with TET2 as a major target for mutations and translocations leading to a wide range of myeloid leukemias, related myelodysplastic conditions (ASXL1 and ASXL2) and the Bohring-Opitz syndrome, a developmental disorder (ASXL1). Most animal ASXL proteins (except the Drosophila Asx protein) contain an N-terminal domain that is also found in several other eukaryotic chromatin proteins (including Arabidopsis HB1), diverse restriction endonucleases and DNA glycosylases, the RNA polymerase delta subunit of Gram-positive bacteria and certain bacterial proteins that combine features of the RNA polymerase α-subunit and sigma factors. This domain adopts the winged helix-turn-helix fold and is predicted to bind DNA. It has been named HARE-type HTH, for HB1, ASXL, restriction endonuclease, based on the proteins in which it was detected [1,2].

The HARE-type HTH domain consists of four α-helices (helices I, II, III, and IV) and an antiparallel β-sheet composed of three short β-strands at the top of a "twisted tripod" formed by helices II, III, and IV (see <PDB:2KRC>) [2].

The profile we developed covers the entire HARE-type HTH domain.

Last update:

January 2020 / First entry.

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Technical section

PROSITE method (with tools and information) covered by this documentation:

HTH_HARE, PS51913; HARE-type HTH domain profile  (MATRIX)


References

1AuthorsAravind L. Iyer L.M.
TitleThe HARE-HTH and associated domains: novel modules in the coordination of epigenetic DNA and protein modifications.
SourceCell. Cycle. 11:119-131(2012).
PubMed ID22186017
DOI10.4161/cc.11.1.18475

2AuthorsMotackova V. Sanderova H. Zidek L. Novacek J. Padrta P. Svenkova A. Korelusova J. Jonak J. Krasny L. Sklenar V.
TitleSolution structure of the N-terminal domain of Bacillus subtilis delta subunit of RNA polymerase and its classification based on structural homologs.
SourceProteins 78:1807-1810(2010).
PubMed ID20310067
DOI10.1002/prot.22708



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